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The locality of Zwartbas is situated at the border of Namibia and South Africa about 15 km west of Noordoewer. The mapped area is confined by the Tandjieskoppe Mountains in the north and the Orange River in the south. Outcropping rocks are predominantly sediments of the Nama Group and of the Karoo Supergroup. During the compilation of this paper doubts arose about the correct classification of the Nama rocks as it is found in literature. Since no certain clues were found to revise the classification of the Nama rocks, the original classification remains still valid. Thus the Kuibis and Schwarzrand Subgroup constitute the Nama succession and date it to Vendian age. A glacial unconformity represents a hiatus for about 260 Ma. This is covered by sediments of the Karoo Supergroup. Late Carboniferous and early Permian glacial deposits of diamictitic shale of the Dwyka and shales of the Ecca Group overlie the unconformity. The shales of the Dwyka Group contain fossiliferous units and volcanic ash-layers. A sill of the Jurassic Tandjiesberg Dolerite Complex (also Karoo Supergroup) intruded rocks at the Dwyka-Ecca-boundary. Finally fluvial and aeolian deposits and calcretes of the Cretaceous to Tertiary Kalahari Group and recent depositionary events cover the older rocks occasionally.
At Zwartbas, about 10 km west of Vioolsdrif, southern Namibia, the Dwyka succession is composed of tillites and distal fossiliferous dropstone-bearing glacio-marine shales. The completely exposed Dwyka succession is interbedded with thin bentonites, altered distal pyroclastic deposits, which were derived from the magmatic arc at the southern rim of Gondwana. Dropstone-bearing and dropstonefree sequences intercalate with four diamictites, of which the two lowest were certainly recognised as tillites. Four events of deglaciation were proven at Zwartbas and thus consist with correlative deposits in southern Africa. Numerous fossilised fishes, trace fossils, and plant fragments appear frequently within the lower half of the Dwyka succession whereas trace fossils were principally found in the complete succession. Although the environmental determination is quite problematic, the fossil assemblage rather implies proximal, shallow water conditions with temporary restricted oxygenation. The hinterland was covered with considerable vegetation, which points to a moderate climate. Water salinity determinations based on shale geochemistry rectify contrary palaeontological results and point to rather brackish or non-marine conditions in comparison to present-day salinites. Geochemical analyses of the bentonites relate the pyroclastic deposits with acid to intermediate source magmas, as they are known from the magmatic arc in present-day Patagonia. Tectono-magmatic comparisons furthermore emphasise a syn-collision or volcanic-arc situation of the magma source. However, significant cyclicity in the production of the pyroclastic deposits was not observed. Radiometric age determinations of two tuff beds clearly date the onset of glacial activity into the Late Carboniferous.
This study aims at reconstructing landscape evolution in the Quebrada de Purmamarca, NW-Argentina. Thorough mapping of the existing landforms and present morphodynamic situation was conducted on the base of intensive field work and the interpretation of remote sensing imagery. Aside from geomorphological mapping, field work focused on the description of numerous sedimentological and pedological profiles. The analysis of these profiles was supported by laboratory data from field samples (granulometry, CaCO3 content) but also by a 14C age date. With particular regard to pedological questions, several samples from soil crust were micromorphologically analysed and interpreted. The resulting data allowed the reconstruction of several phases of landscape evolution in the Quebrada de Purmamarca back to the Miocene. During this phase, the Andes were still a landscape of relatively low relief being subject to processes of planation under conditions markedly more humid than today. Highly faulted and deformed fanglomerates are the first evidence of a progressing uplift coupled under an increasingly arid climate. As a consequence of continued uplift and alternating phases of erosion and aggradation, large terrace systems have formed. Particularly the youngest terrace level shows good preservation. Against the background of the intense climatic changes characteristic for the Pleistocene, these terraces have been the major focus of this study. They are built up almost entirely from coarse debris-flow sediments which are thought to be the result of a significant drop of the periglacial belt of more than 1,000 meters. This interpretation is confirmed by a variety of relict periglacial landforms like “glatthang” morphology (smooth topography), sheets of frost debris and asymmetric valleys. As sediment supply from periglacial debris production exceeded the transport capacity of the drainage system leading to the dominance of depositional processes. Aggradation has been interrupted or at least weakened several times as reflected by two lacustrine to fluvial intervals within the terrace deposits. In this context, particularly the younger interval might announce a shift in morphodynamics around 49 ka BP (14C age), when the phase of terrace aggradation grades into a phase of dominant alluvial fan activity. On the terrace surfaces a well-developed reddish soil has developed. It is interpreted to indicate a phase of increased humidity possibly in relation with the “Minchin” wet phase between 40 ka BP and 25 ka BP. At many places, this reddish soil is overlain by a markedly cemented sand crust. Based on the good sorting of medium and fine sand, this sand crust could be interpreted as fluvio-eolian sediment. Its deposition under very arid and cold climatic conditions may be attributed to the Late Glacial Maximum (LGM). However, the sand crust shows signs of erosion at many places and has not been observed anywhere below the level of the terrace surface. Therefore the onset of severe erosion and incision resulting in the evacuation of enormous quantities of sediment from the study area is assumed to postdate the LGM, possibly due to increased discharge rates during a wetter Lateglacial. Regardless of its timing, the intense incision is likely to have cut down to below the present floodplain evidently causing several mass wasting events in the study area. Since the early Holocene a number of short-term changes seem to have been responsible for the landscape evolution of the Quebrada de Purmamarca. More humid phases of pronounced slope smoothing have alternated with semi-arid phases of longer duration. The well-developed, polycyclic calcretes on top of the inactive terraces and alluvial fans give evidence for these changes. At present, the marked desert pavement on top of most terraces and alluvial fan surfaces prevent soil. The concentration of runoff on these pavements amplifies badland formation and alluvial fan activity along the terrace slopes. The presently observed floodplain aggradation may be attributed to these processes but considering the severe gullying reaching far into the upper study area, the aggradation may as well reflect a more general and regional trend.
Participants trained aiming movements of the right hand to several targets with a prism-like virtual displacement of the location of one of the targets, receiving either terminal or continuous visual feedback. After training, the same targets were to be reached with the untrained left hand under manipulated feedback conditions. The right hand movements continuously adapted to the unnoticed visual displacement, significantly less with continuous than with terminal feedback. Under terminal but not under continuous feedback the adaptation to the manipulated target generalized to targets in the same horizontal direction but not to targets in the opposite direction. Finally, the movements of the untrained left hand showed the same qualitative changes to the targets as the movements of the trained right hand. The data are in line with the notion that the adaptation of the right hand movements is mainly based on a re-interpretation of target locations on which movement control of both hands draws.
Burn severity was measured within the Mediterranean sclerophyll forests of south-west Western Australia (WA) using remote sensing data from the Moderate Resolution Imaging Spectroradiometer (MODIS). The region of south-west WA is considered as a high fire prone landscape and is managed by the state government’s Department of Conservation and Land Management (CALM). Prescribed fuel reduction burning is used as a management tool in this region. The measurement of burn severity with remote sensing data focused on monitoring the success and impact of prescribed burning and wildfire in this environment. The high temporal resolution of MODIS with twice daily overpasses in this area was considered highly favourable, as opportunities for prescribed burning are temporally limited by climatic conditions. The Normalised Burn Ratio (NBR) was investigated to measure burn severity in the forested area of south-west WA. This index has its heritage based on data from the Landsat TM/ETM+ sensors (Key and Benson, 1999 [1],[2]) and was transferred from Landsat to MODIS data. The measurement principally addresses the biomass consumption due to fire, whereas the change detected between the pre-fire image and the post-fire image is quantified by the ÄNBR. The NBR and the Normalised Difference Vegetation Index (NDVI) have been applied to MODIS and Landsat TM/ETM+ data. The spectral properties and the index values of the remote sensing data have been analysed within different burnt areas. The influence of atmospheric and BRDF effects on MODIS data has been investigated by comparing uncorrected top of atmosphere reflectance and atmospheric and BRDF corrected reflectance. The definition of burn severity classes has been established in a field trip to the study area. However, heterogeneous fire behaviour and patchy distribution of different vegetation structure made field classification difficult. Ground truth data has been collected in two different types of vegetation structure present in the burnt area. The burn severity measurement of high resolution Landsat data was assessed based on ground truth data. However, field data was not sufficient for rigorous validation of remote sensing data. The NBR index images of both sensors have been calibrated based on training areas in the high resolution Landsat image. The burn severity classifications of both sensors are comparable, which demonstrates the feasibility of a burn severity measurement using moderate spatial resolution 250m MODIS data. The normalisation through index calculation reduced atmospheric and BRDF effects, and thus MODIS top of at-mosphere data has been considered suitable for the burn severity measurement. The NBR could not be uniformly applied, as different structures of vegetation influenced the range of index values. Furthermore, the index was sensitive to variability in moisture content. However, the study concluded that the NBR on MODIS data is a useful measure of burn severity in the forested area of south-west WA.
The internal transcribed spacer 2 (ITS2) of the ribosomal gene repeat is an increasingly important phylogenetic marker whose RNA secondary structure is widely conserved across eukaryotic organisms. The ITS2 database aims to be a comprehensive resource on ITS2 sequence and secondary structure, based on direct thermodynamic as well as homology modelled RNA folds. Results: (a) A rebuild of the original ITS2 database generation scripts applied to a current NCBI dataset reveal more than 60,000 ITS2 structures. This more than doubles the contents of the original database and triples it when including partial structures. (b) The end-user interface was rewritten, extended and now features user-defined homology modelling. (c) Other possible RNA structure discovery methods (namely suboptimal and shape folding) prove helpful but are not able to replace homology modelling. (d) A use case of the ITS2 database in conjunction with other tools developed at the department gave insight into molecular phylogenetic analysis with ITS2.
The scope of this work is to develop a novel single-molecule imaging technique by combining atomic force microscopy (AFM) and optical fluorescence microscopy. The technique is used for characterizing the structural properties of multi-protein complexes. The high-resolution fluorescence microscopy and AFM are combined (FIONA-AFM) to allow for the identification of individual proteins in such complexes. This is achieved by labeling single proteins with fluorescent dyes and determining the positions of these fluorophores with high precision in an optical image. The same area of the sample is subsequently scanned by AFM. Finally, the two images are aligned and the positions of the fluorophores are displayed on top of the topographical data. Using quantum dots as fiducial markers in addition to fluorescently labeled proteins, fluorescence and AFM information can be aligned with an accuracy better than 10 nm, which is sufficient to identify single fluorescently labeled proteins in most multi-protein complexes. The limitations of localization precision and accuracy in fluorescence and AFM images are investigated, including their effects on the overall registration accuracy of FIONA-AFM hybrid images. This combination of the two complementary techniques opens a wide spectrum of possible applications to the study of protein interactions, because AFM can yield high resolution (5–10 nm) information about the conformational properties of multi-protein complexes while the fluorescence can indicate spatial relationships of the proteins within the complexes. Additionally, computer simulations are performed in order to validate the accuracy of the registration algorithm.
From the introduction: It is not always easy to define what a word means. We can choose between a variety of possibilities, from simply pointing at the correct object as we say its name to lengthy definitions in encyclopaedias, which can sometimes fill multiple pages. Although the former approach is pretty straightforward and is also very important for first language acquisition, it is obviously not a practical solution for defining the semantics of the whole lexicon. The latter approach is more widely accepted in this context, but it turns out that defining dictionary and encyclopaedia entries is not an easy task. In order to simplify the challenge of defining the meaning of words, it is of great advantage to organize the lexicon in a way that the structure in which the words are integrated gives us information about the meaning of the words by showing their relation to other words. These semantic relations are the focal point of this paper. In the first chapter, different ways to describe meaning will be discussed. It will become obvious why semantic relations are a very good instrument to organizing the lexicon. The second chapter deals with WordNet, an electronic lexical database which follows precisely this approach. We will examine the semantic relations which are used in WordNet and we will study the distinct characteristics of each of them. Furthermore, we will see which contribution is made by which relation to the organization of the lexicon. Finally, we will look at the downside of the fact that WordNet is a manually engineered network by examining the shortcomings of WordNet. In the third chapter, an alternative approach to linguistics is introduced. We will discuss the principles of corpus linguistics and, using the example of the British National Corpus, we will consider possibilities to extract semantic relations from language corpora which could help to overcome the deficiencies of the knowledge based approach. In the fourth chapter, I will describe a project the goal of which is to extend WordNet by findings from cognitive linguistics. Therefore, I will discuss the development process of a piece of software that has been programmed in the course of this thesis. Furthermore, the results from a small‐scale study using this software will be analysed and evaluated in order to check for the success of the project.
The capabilities of a new approach towards the foundations of Statistical Mechanics are explored. The approach is genuine quantum in the sense that statistical behavior is a consequence of objective quantum uncertainties due to entanglement and uncertainty relations. No additional randomness is added by hand and no assumptions about a priori probabilities are made, instead measure concentration results are used to justify the methods of Statistical Physics. The approach explains the applicability of the microcanonical and canonical ensemble and the tendency to equilibrate in a natural way. This work contains a pedagogical review of the existing literature and some new results. The most important of which are: i) A measure theoretic justification for the microcanonical ensemble. ii) Bounds on the subsystem equilibration time. iii) A proof that a generic weak interaction causes decoherence in the energy eigenbasis. iv) A proof of a quantum H-Theorem. v) New estimates of the average effective dimension for initial product states and states from the mean energy ensemble. vi) A proof that time and ensemble averages of observables are typically close to each other. vii) A bound on the fluctuations of the purity of a system coupled to a bath.
Bees are subject to permanent threat from predators such as ants. Their nests with large quantities of brood, pollen and honey represent lucrative targets for attacks whereas foragers have to face rivalry at food sources. This thesis focused on the role of stingless bees as third party interactor on ant-aphid-associations as well as on the predatory potential represented by ants and defense mechanisms against this threat. Regular observations of an aphid infested Podocarpus for approaching stingless bees yielded no results. Another aim of this thesis was the observation of foraging habits of four native and one introduced ant species for assessment of their predatory potential to stingless bees. All species turned out to be dietary balanced generalists with one mostly carnivorous species and four species predominantly collecting nectar roughly according to optimal foraging theory. Two of the species monitored, Rhytidoponera metallica and Iridomyrmex rufoniger were considered potential nest robbers. As the name implies, stingless bees lack the powerful weapon of their distant relatives; hence they specialized on other defense strategies. Resin is an important, multipurpose resource for stingless bees that is used as material for nest construction, antibiotic and for defensive means. For the latter purpose highly viscous resin is either directly used to stick down aggressors or its terpenic compounds are included in the bees cuticular surface. In a feeding choice experiment, three ant species were confronted with the choice between two native bee species - Tetragonula carbonaria and Austroplebeia australis - with different cuticular profiles and resin collection habits. Two of the ant species, especially the introduced Tetramorium bicarinatum did not show any preferences. The carnivorous R. metallica predominantly took the less resinous A. australis as prey. The reluctance towards T. carbonaria disappeared when the resinous compounds on its cuticle had been washed off with hexane. To test whether the repulsive reactions were related to the stickiness of the resinous surface or to chemical substances, hexane extracts of bees’ cuticles, propolis and three natural tree resins were prepared. In the following assay responses of ants towards extract treated surfaces were observed. Except for one of the resin extracts, all tested substances had repellent effects to the ants. Efficacy varied with the type of extract and species. Especially to the introduced T. bicarinatum the cuticular extract had no effect. GCMS-analyses showed that some of the resinous compounds were also found in the cuticular profile of T. carbonaria which featured reasonable analogies to the resin of Corymbia torelliana that is highly attractive for stingless bees. The results showed that repellent effects were only partially related to the sticky quality of resin but were rather caused by chemical substances, presumably sesqui- and diterpenes. Despite its efficacy this defense strategy only provides short time repellent effects sufficient for escape and warning of nest mates to initiate further preventive measures.